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- Haldan_Keffer_Hartline abstract "Haldan Keffer Hartline ForMemRS (December 22, 1903 – March 17, 1983) was an American physiologist who was a co-recipient (with George Wald and Ragnar Granit) of the 1967 Nobel Prize in Physiology or Medicine for his work in analyzing the neurophysiological mechanisms of vision.Hartline received his undergraduate education from Lafayette College in Easton, Pennsylvania, graduating in 1923. He began his study of retinal electrophysiology as a National Research Council Fellow at Johns Hopkins University, Baltimore, receiving his M.D. in 1927. After attending the universities of Leipzig and Munich as an Eldridge Johnson traveling research scholar from the University of Pennsylvania, he returned to the US to take a position in the Eldridge Reeves Johnson Foundation for Medical Physics at Penn, which was under the directorship of Detlev W. Bronk at that time. In 1940–1941, he was Associate Professor of Physiology at Cornell Medical College in New York City, but returned to Penn and stayed until 1949. Then he became professor of biophysics and chairman of the department at Johns Hopkins in 1949. One of Hartline's graduate students at Johns Hopkins, Paul Greengard, who also won the Nobel Prize. Hartline joined the staff of Rockefeller University, New York City, in 1953 as professor of neurophysiology.Hartline investigated the electrical responses of the retinas of certain arthropods, vertebrates, and mollusks, because their visual systems are much simpler than those of humans and thus easier to study. He concentrated his studies on the eye of the horseshoe crab (Limulus polyphemus). Using minute electrodes, he obtained the first record of the electrical impulses sent by a single optic nerve fibre when the receptors connected to it are stimulated by light. He found that the photoreceptor cells in the eye are interconnected in such a way that when one is stimulated, others nearby are depressed, thus enhancing the contrast in light patterns and sharpening the perception of shapes. Hartline thus built up a detailed understanding of the workings of individual photoreceptors and nerve fibres in the retina, and he showed how simple retinal mechanisms constitute vital steps in the integration of visual information.".
- Haldan_Keffer_Hartline almaMater Johns_Hopkins_School_of_Medicine.
- Haldan_Keffer_Hartline almaMater Lafayette_College.
- Haldan_Keffer_Hartline award Nobel_Prize_in_Physiology_or_Medicine.
- Haldan_Keffer_Hartline birthDate "1903-12-22".
- Haldan_Keffer_Hartline birthPlace Bloomsburg,_Pennsylvania.
- Haldan_Keffer_Hartline deathDate "1983-03-17".
- Haldan_Keffer_Hartline deathPlace Fallston,_Maryland.
- Haldan_Keffer_Hartline doctoralAdvisor August_Herman_Pfund.
- Haldan_Keffer_Hartline field Physiology.
- Haldan_Keffer_Hartline knownFor Visual_perception.
- Haldan_Keffer_Hartline thumbnail Haldan_Keffer_Hartline_nobel.jpg?width=300.
- Haldan_Keffer_Hartline wikiPageExternalLink hartline-bio.html.
- Haldan_Keffer_Hartline wikiPageExternalLink hartline-haldan.pdf.
- Haldan_Keffer_Hartline wikiPageID "591723".
- Haldan_Keffer_Hartline wikiPageLength "7290".
- Haldan_Keffer_Hartline wikiPageOutDegree "44".
- Haldan_Keffer_Hartline wikiPageRevisionID "703223866".
- Haldan_Keffer_Hartline wikiPageWikiLink Arthropod.
- Haldan_Keffer_Hartline wikiPageWikiLink Atlantic_horseshoe_crab.
- Haldan_Keffer_Hartline wikiPageWikiLink August_Herman_Pfund.
- Haldan_Keffer_Hartline wikiPageWikiLink Baltimore.
- Haldan_Keffer_Hartline wikiPageWikiLink Biophysics.
- Haldan_Keffer_Hartline wikiPageWikiLink Bloomsburg,_Pennsylvania.
- Haldan_Keffer_Hartline wikiPageWikiLink Category:1903_births.
- Haldan_Keffer_Hartline wikiPageWikiLink Category:1983_deaths.
- Haldan_Keffer_Hartline wikiPageWikiLink Category:American_Nobel_laureates.
- Haldan_Keffer_Hartline wikiPageWikiLink Category:American_neuroscientists.
- Haldan_Keffer_Hartline wikiPageWikiLink Category:Foreign_Members_of_the_Royal_Society.
- Haldan_Keffer_Hartline wikiPageWikiLink Category:Johns_Hopkins_University_alumni.
- Haldan_Keffer_Hartline wikiPageWikiLink Category:Medical_physicists.
- Haldan_Keffer_Hartline wikiPageWikiLink Category:Members_of_the_United_States_National_Academy_of_Sciences.
- Haldan_Keffer_Hartline wikiPageWikiLink Category:Nobel_laureates_in_Physiology_or_Medicine.
- Haldan_Keffer_Hartline wikiPageWikiLink Detlev_Bronk.
- Haldan_Keffer_Hartline wikiPageWikiLink Fallston,_Maryland.
- Haldan_Keffer_Hartline wikiPageWikiLink George_Wald.
- Haldan_Keffer_Hartline wikiPageWikiLink Horseshoe_crab.
- Haldan_Keffer_Hartline wikiPageWikiLink Johns_Hopkins_School_of_Medicine.
- Haldan_Keffer_Hartline wikiPageWikiLink Johns_Hopkins_University.
- Haldan_Keffer_Hartline wikiPageWikiLink Lafayette_College.
- Haldan_Keffer_Hartline wikiPageWikiLink Lateral_inhibition.
- Haldan_Keffer_Hartline wikiPageWikiLink Leipzig_University.
- Haldan_Keffer_Hartline wikiPageWikiLink Ludwig_Maximilian_University_of_Munich.
- Haldan_Keffer_Hartline wikiPageWikiLink Mollusca.
- Haldan_Keffer_Hartline wikiPageWikiLink Nobel_Prize_in_Physiology_or_Medicine.
- Haldan_Keffer_Hartline wikiPageWikiLink Optic_nerve.
- Haldan_Keffer_Hartline wikiPageWikiLink Paul_Greengard.
- Haldan_Keffer_Hartline wikiPageWikiLink Photoreceptor_cell.
- Haldan_Keffer_Hartline wikiPageWikiLink Physiology.
- Haldan_Keffer_Hartline wikiPageWikiLink Ragnar_Granit.
- Haldan_Keffer_Hartline wikiPageWikiLink Retina.
- Haldan_Keffer_Hartline wikiPageWikiLink Rockefeller_University.
- Haldan_Keffer_Hartline wikiPageWikiLink Royal_Society.
- Haldan_Keffer_Hartline wikiPageWikiLink University_of_Pennsylvania.
- Haldan_Keffer_Hartline wikiPageWikiLink Vertebrate.
- Haldan_Keffer_Hartline wikiPageWikiLink Visual_perception.
- Haldan_Keffer_Hartline wikiPageWikiLink Visual_system.
- Haldan_Keffer_Hartline wikiPageWikiLinkText "H. Keffer Hartline".
- Haldan_Keffer_Hartline wikiPageWikiLinkText "Haldan Keffer Hartline".
- Haldan_Keffer_Hartline wikiPageWikiLinkText "Hartline".
- Haldan_Keffer_Hartline wikiPageWikiLinkText "Hartline, Haldan Keffer".
- Haldan_Keffer_Hartline almaMater Johns_Hopkins_School_of_Medicine.
- Haldan_Keffer_Hartline almaMater Lafayette_College.
- Haldan_Keffer_Hartline birthDate "1903-12-22".
- Haldan_Keffer_Hartline birthPlace "Bloomsburg, Pennsylvania, US".
- Haldan_Keffer_Hartline caption "Haldan Keffer Hartline, circa 1958".
- Haldan_Keffer_Hartline deathDate "1983-03-17".
- Haldan_Keffer_Hartline deathPlace "Fallston, Maryland, US".
- Haldan_Keffer_Hartline doctoralAdvisor August_Herman_Pfund.
- Haldan_Keffer_Hartline field Physiology.
- Haldan_Keffer_Hartline knownFor Visual_perception.
- Haldan_Keffer_Hartline name "Haldan Keffer Hartline".
- Haldan_Keffer_Hartline nationality "American".
- Haldan_Keffer_Hartline prizes Nobel_Prize_in_Physiology_or_Medicine.
- Haldan_Keffer_Hartline wikiPageUsesTemplate Template:Authority_control.
- Haldan_Keffer_Hartline wikiPageUsesTemplate Template:Birth_date.
- Haldan_Keffer_Hartline wikiPageUsesTemplate Template:Citation_needed.
- Haldan_Keffer_Hartline wikiPageUsesTemplate Template:Death_date_and_age.
- Haldan_Keffer_Hartline wikiPageUsesTemplate Template:Infobox_scientist.
- Haldan_Keffer_Hartline wikiPageUsesTemplate Template:Nobel_Prize_in_Physiology_or_Medicine_Laureates_1951-1975.
- Haldan_Keffer_Hartline wikiPageUsesTemplate Template:Post-nominals.
- Haldan_Keffer_Hartline wikiPageUsesTemplate Template:Reflist.
- Haldan_Keffer_Hartline workInstitutions Johns_Hopkins_University.
- Haldan_Keffer_Hartline workInstitutions University_of_Pennsylvania.
- Haldan_Keffer_Hartline subject Category:1903_births.
- Haldan_Keffer_Hartline subject Category:1983_deaths.
- Haldan_Keffer_Hartline subject Category:American_Nobel_laureates.
- Haldan_Keffer_Hartline subject Category:American_neuroscientists.
- Haldan_Keffer_Hartline subject Category:Foreign_Members_of_the_Royal_Society.
- Haldan_Keffer_Hartline subject Category:Johns_Hopkins_University_alumni.
- Haldan_Keffer_Hartline subject Category:Medical_physicists.
- Haldan_Keffer_Hartline subject Category:Members_of_the_United_States_National_Academy_of_Sciences.
- Haldan_Keffer_Hartline subject Category:Nobel_laureates_in_Physiology_or_Medicine.
- Haldan_Keffer_Hartline hypernym Physiologist.
- Haldan_Keffer_Hartline type Agent.
- Haldan_Keffer_Hartline type Award.
- Haldan_Keffer_Hartline type Biologist.
- Haldan_Keffer_Hartline type Person.
- Haldan_Keffer_Hartline type Scientist.
- Haldan_Keffer_Hartline type Person.